Institute of Vertebrate Paleontology and Paleoanthropology
Chinese Academy of Sciences
Institute of Vertebrate Paleontology and Paleoanthropology
Chinese Academy of Sciences
Faculty and Staff
WANG Yuan
WANG Yuan

Title: Professor

Fax: 010-68337001

E-mail: wangyuan@ivpp.ac.cn

Education and Appointments
  • 2008~present  Researcher, IVPP, Chinese Academy of Sciences
  • 2004~present  Director, Paleozoological Museum of China
  • 2002  PhD, Chinese Academy of Sciences, Paleontology and Stratigraphy
  • 1996  MA, University of Kansas, Systematics
  • 1994  MSc, Chinese Academy of Sciences, Paleontology and Stratigraphy
  • 1991  BSc, Peking University, Paleontology and Stratigraphy
Research Interests
My research interests focus on the evolution of fossil amphibians and small reptiles from China, on their morphological evolution and ontogenetic development (particularly osteological), systematics and phylogeny, and paleobiogeographic reconstruction. Current research is mainly on the well-preserved and mostly articulated frogs, salamanders, and lizards of the Early Cretaceous Jehol Biota and of associated earlier horizons (e.g. Jurassic Daohugou Bed). The fossil localities are mainly in northeastern China (Liaoning, Hebei Provinces and Inner Mongolia Autonomous Region). Other on-going work includes lizards and temnospondyl amphibians from the Jurassic of Xinjiang Autonomous Region and frogs from Cenozoic localities in eastern and southern China.
Public Services
Honors
Selected Publications

Selected Publications of WANG Yuan

* denotes SCI journals; 

2021

1.       *Rong Y F, Vasilyan D, Dong L P, Wang Y, 2020,2021. Revision of Chunerpeton tianyiense (Lissamphibia, Caudata): Is it a cryptobranchid salamander? Palaeoworld, https://doi.org/10.1016/j.palwor.2020.12.001. (16pp) [PDF]

2.       *Li L, Zhang J L, Wang X L, Wang Y, Tong H Y, 2021. First turtle remains from the Middle-Late Jurassic Yanliao Biota, NE China. Science Technology and Engineering Journal, 7(1): 1~11. [PDF]

2.       *Li L, Zhang J L, Wang X L, Wang Y, Tong H Y, 2021. First turtle remains from the Middle-Late Jurassic Yanliao Biota, NE China. Science Technology and Engineering Journal, 7(1): 1~11. [PDF]

 

2020

1.       *Dong L P, Matsumoto R, Kusuhashi N, Wang Y Q, Wang Y, Evans S E, 2020. A new choristodere (Reptilia, Choristodera) from an Aptian–Albian coal deposit in China. Journal of Systematic Paleontology. http://dx.doi.org/10.1080/14772019.2020.1749147. 20pp. [pdf]

2.       *Rong Y F, Vasilyan D, Dong L P, Wang Y (in press). Revision of Chunerpeton tianyiense (Lissamphibia, Caudata): is it a cryptobranchid salamander? Palaeoworld.

3.       Xu X, Zhou Z, Wang Y, Wang M. 2020. Study on the Jehol Biota: Recent advances and future prospects. Science China: Earth Sciences, 63:757~773. https://doi.org/10.1007/s11430-019-9509-3. [PDF]

 

2019

4.       Wang Y, Wu F X, Jin H Y, Ge X, Yang J, 2019. 90 Fossils Attest to 90 Years of Glory. Beijing: Popular Science Press. 245pp. [in Chinese] [pdf]

5.       *Matsumoto R, Dong L P, Wang Y, Evans S E, 2019. The first record of a nearly complete choristodere (Reptilia: Diapsida) from the Upper Jurassic of Hebei Province, People’s Republic of China. Journal of Systematic Palaeontology, 17(12): 1031~1048, DOI: 10.1080/14772019.2018.1494220.

6.       *Dong L P, Wang Y, Mou L J, Zhang G Z, Evans S E, 2019. A new Jurassic lizard from China. in Steyer J.-S., Augé M. L. & Métais G. (eds), Memorial Jean-Claude Rage: A life of paleo-herpetologist. Geodiversitas 41 (16): 623~641. https://doi.org/10.5252/geodiversitas2019v41a16.

 

2018

7.       Wang Y, Ge X, Xing L D, Xie D, Ma N, 2018. Listening to the Stories of Fossils. Beijing: Popular Science Press. 271pp. [in Chinese] [pdf]

8.       *Dong L P, Xu X, Wang Y, Evans S E, 2018. The Lizard Genera Bainguis and Parmeosaurus from the Upper Cretaceous of China and Mongolia. Cretaceous Research 85, 95~108. [PDF]

 

2017

9.       *Dong L P, Wang Y, Evans S E, 2017. A new lizard (Reptilia: Squamata) from the Lower Cretaceous Yixian Formation of China, with a taxonomic revision of Yabeinosaurus, Cretaceous Research, 72: 161~171. (doi: 10.1016/j.cretres.2016.12.017, published on-line in 2016). [PDF]

10.    Sullivan C, Wang Y, Choo B, 2017. From Fish to Human: the March of Vertebrate Life in China. Beijing: Popular Science Press. 307pp. [Collector’s edition in Chinese] [pdf]

11.    Xu X, Zhou Z H, Sullivan C, and Wang Y, 2017. The Yanliao Biota: a trove of exceptionally preserved Middle-Late Jurassic terrestrial life forms. In: Fraser N C and Sues H-D (eds). Terrestrial Conservation Lagerst?tten: Windows into the Evolution of Life on Land. Edinburgh & London: Dunedin Academic Press. 131~167.

12.    Zhou Z H, Wang Y, Xu X, Ren D, 2017. The Jehol Biota: an exceptional window into the Early Cretaceous terrestrial ecosystem. In: Fraser N C and Sues H-D (eds). Terrestrial Conservation Lagerst?tten: Windows into the Evolution of Life on Land. Edinburgh & London: Dunedin Academic Press. 169~214.

13.    *Zhou Z H, Wang Y, 2017. Vertebrate assemblages of the Jurassic Yanliao Biota and the Early Cretaceous Jehol Biota: Comparisons and implications. Palaeoworld, 26(2): 241~252 (http://dx.doi.org/10.1016/j.palwor.2017.01.002) [PDF]

14.    *Wang Y, Ro?ek Z, Dong L P, 2017. A new pelobatoid frog from the lower Eocene of southern China. Palaeobiodiversity and Palaeoenvironments (DOI: 10.1007/s12549-017-0304-1). 18pp. [PDF]

 

2016

15.    Han F L, Wang Y, Sullivan C, Wang Y Q, Qin Z C, Xu X, 2016. Chapter 7: The Vertebrates. In: Huang D Y (ed.), The Daohugou Biota. Shanghai: Shanghai Scientific and Technical Publisher. 218~251 [in Chinese]. [pdf]

16.    *Wang Y, Dong L P, Evans S E, 2016. Polydactyly and other limb abnormalities in the Jurassic salamander Chunerpeton from China. Palaeobiodiversity and Palaeoenvironment (2016) 96:49~59 (DOI 10.1007/s12549-015-0219-7, published on line in 2015). [PDF]

17.    Dong L P, Evans S E, Wang Y, 2016. Taxonomic revision of lizards from the Paleocene deposits of the Qianshan Basin, Anhui, China. Vertebrata Palasiatica, 54(3): 243~268. [PDF]

18.    *Xu X, Zhou Z H, Sullivan C, Wang Y, Ren D, 2016. An Updated Review of the Middle-Late Jurassic Yanliao Biota: Chronology, Taphonomy, Paleontology, and Paleoecology. Acta Geologica Sinica (English Edition), 90 (6): 2229~2243. [PDF]

 

2015

19.    Wang Y et al., 2015. Amphibians (Fascicle 1, Serial no. 5). In: Editorial Committee of Palaeovertebrata Sinica (ed.): Palaeovertebrata Sinica (Volume II: Amphibians, Reptiles and Avians). Science Press. 143pp. [in Chinese]

20.    Sullivan C, Wang Y, Choo B, 2015. From Fish to Human: the March of Vertebrate Life in China. Beijing: Popular Science Press. 546pp. [in English and Chinese]

 

2014

21.    *Sullivan C, Wang Y, Hone D W E, Wang Y Q, Xu X, Zhang F C, 2014. The Vertebrates of the Jurassic Daohugou Biota of Northeastern China. Journal of Vertebrate Paleontology. 34(1):1~37. [PDF]

 

2013

22.    Zhou Z H (ed.), Wang X D, Wang Y (co-eds.), 2013. 100,000 Whys (Palaeontology). Shanghai, Juvenile and Children’s Publishing House. 201pp. [in Chinese]

23.    Wang Y, 2013. The stories behind two status. In: Wang Q, 2013. From Crystal Palace to the Darwin Centre: An Architectural Evolution of the Natural Histroy Museum in London. Beijing: China Architecture Industry Press. 108~114. [in Chinese] [pdf]

24.    *Dong L, Roek Z, Wang Y, Jones M E H, 2013. Anurans from the Lower Cretaceous Jehol Group of Western Liaoning, China. PLoS ONE 8(7): e69723. doi:10.1371/journal.pone.0069723. [PDF]

 

2012

25.    *Rocek Z, Wang Y, Dong L, 2012. Post-metamorphic development of Early Cretaceous frogs as a tool for taxonomic comparisons. Journal of Vertebrate Paleontology, 32(6): 1285~1292. [PDF]

26.    *Evans S, Wang Y, 2012. New material of the Early Cretaceous lizard Yabeinosaurus from China. Cretaceous Research, 34: 48~60 (doi:10.1016/j.cretres.2011.10.004: published on line in 2011). [PDF]

27.    Dong L P, Huang D Y, Wang Y, 2012. Two Jurassic salamanders with stomach contents from Inner Mongolia, China. Chinese Sci Bull, 57, doi:10.1007/s11434-011-4729-z. [PDF]

 

2011

28.    *Rocek Z, Dong L, Pikryl T, Sun C, Tan J, Wang Y, 2011. Fossil frogs (Anura) from Shanwang (Middle Miocene; Shandong Province, China). Geobios, 44(5): 499~518 (doi:10.1016/j.geobios.2010.11.009) [PDF]

29.    *Wang Y, Evans S E, 2011. A gravid lizard from the Cretaceous of China and the early history of squamate viviparity. Naturwissenschaften, 98(9): 739~743. (DOI:10.1007/s00114-011-0820-1) [PDF]

 

2010

30.    Zhou Z H, Jin F, Wang Y, 2010. Vertebrate assemblages from the Middle-Late Jurassic Yanliao Biota in Northeast China. Earth Science Frontiers, 17(special issue): 252~254. [pdf]

31.    *Zhou Z H, Wang Y, 2010. Major features of the vertebrate diversity of the Early Cretaceous Jehol Biota and their paleoecological implications. Journal of Earth Science, 21(special issue): 228~230. [pdf]

32.    Wang Y, Dong L P, Evans S E, 2010. Jurassic-Cretaceous herpetofaunas from the Jehol associated strata in NE China: evolutionary and ecological implications. Bulletin of the Chinese Academy of Sciences, 24(2): 76~79. [PDF]

33.    *Evans S E, Wang Y, 2010. A new lizard (Reptilia: Squamata) with exquisite preservation of soft tissue from the Lower Cretaceous of Inner Mongolia, China. Journal of Systematic Palaeontology, 8(1): 81~95. [PDF]

34.    *Zhou Z H, Wang Y, 2010. Vertebrate diversity of the Jehol Biota as compared with other Lagerst?tten. Science China (Earth Sciences), 53(12): 1894~1907. [PDF]

35.    Zhou Z H, Wang Y Q, Zhu M, Wang Y, Xu X, Li C, Wang X L., Deng T, 2010. Report on Advances in Veretebrate Paleontology. In: China Association for Science and Technology (ed.), Palaeontological Society of China (compiled), 2009-2010 Report on Advances in Palaeontology. Beijing: China Science and Technology Press. 198~200.

 

2009

36.    Wang Y, 2009. Amphibians. In: The Authorizing Committee for Chinese Terms in Paleontology, Chinese Terms in Paleontology (2nd edition). Beijing: Science Press. 82~85.

37.    Evans S E, Wang Y, 2009. A long-limbed lizard from the Upper Jurassic/Lower Cretaceous of Daohugou, Ningcheng, Nei Mongol, China. Vertebrata PalAsiatica, 47(1): 21~34.

38.    Wang Y, Dong L P, 2009. The evolution of archaic amphibians from China: A long and winding history. In: Sha J G (ed.): A Great Leap in a Century: the Glory of Chinese Palaeontology. Beijing: China Science Press. 238~258. [in Chinese] [pdf]

 

2008

39.    Wang Y, Zhang G L, Sun A L, 2008. Amphibia. In: Li J L, Wu X C, Zhang F C (eds), The Chinese Fossil Reptiles and Their Kin. Beijing: Science Press. 3~25.

40.    Wang Y, Li J L, 2008. Squamata. In: Li J L, Wu X C, Zhang F C (eds), The Chinese Fossil Reptiles and Their Kin. Beijing: Science Press. 115~137.

41.    *Zhang G L, Wang Y, Jones M E H, Evans S E, 2008. A new Early Cretaceous salamander (Regalerpeton weichangensis gen. et sp. nov.) from the Huajiying Formation of northeastern China. Cretaceous Research, doi: 10.1016/j.cretres.2008.10.004. [PDF] /*Zhang G L, Wang Y, Jones M E H, Evans S E, 2009. A new Early Cretaceous salamander (Regalerpeton weichangensis gen. et sp. nov.) from the Huajiying Formation of northeastern China. Cretaceous Research, 30(3): 551~558. [PDF]

 

2007

42.    *Wang Y, Jones M E H, Evans S E, 2007. A juvenile anuran from the Lower Cretaceous Jiufotang Formation, Liaoning, China. Cretaceous Research, 28(2): 235~244. [PDF]

43.    *Evans S E, Wang Y, 2007. A juvenile lizard specimen with well-preserved skin impressions from the Upper Jurassic/Lower Cretaceous of Daohugou, Inner Mongolia, China. Naturwissenschaften, 94(6):431~439. [PDF]

44.    *Evans S E, Wang Y, Jones M E H, 2007. An aggregation of lizard skeletons from the Lower Cretaceous of China. Senckenbergiana Lethaea, 87(1): 109~118. [PDF]

 

2006

45.    *Wang Y, Evans S E, 2006a. A new short-bodied salamander from the Upper Jurassic/Lower Cretaceous of China. Acta Palaeontologica Polonica, 51(1):127~130. [PDF]

46.    Wang Y, Evans S E, 2006b. Advances in the study of fossil amphibians and squamates from China: The past fifteen years. Vertebrata PalAsiatica, 44(1): 60~73. // 王原, 苏珊·E·埃文斯, 2006a. 中国化石两栖类和有鳞类研究进展: 过去15年回顾. 古脊椎动物学报, 44(1): 60~73 (In English with Chinese summary). [PDF]

47.    Wang Y, 2006. Phylogeny and early radiation of Mesozoic lissamphibians from East Asia. In: Rong J Y, Fang Z J, Zhou Z H, Zhan R B, Wang X D, Yuan X L (eds), Originations, Radiations and Biodiversity Changes---Evidence from the Chinese Fossil Record. Beijing: Science Press. 643~663 (in Chinese), 931~936.

 

2005

48.    *Liu J, Wang Y, 2005. The first complete mastodonsauroid skull from the Triassic of China: Yuanansuchus laticeps gen. et sp. nov. Journal of Vertebrate Paleontology, 25(3): 725~728. [PDF]

49.    *Wang Y, Rose C, 2005. Jeholotriton paradoxus (Amphibia: Caudata) from the Lower Cretaceous of southeastern Inner Mongolia, China. Journal of Vertebrate Paleontology, 25(3): 523~532. [PDF]

50.    *Evans S E, Wang Y, Li C, 2005. The Early Cretaceous lizard genus Yabeinosaurus from China: resolving an enigma. Journal of Systematic Palaeontology, 3(4): 319~335. [PDF]

51.    *Evans S E, Wang Y, 2005. The Early Cretaceous lizard Dalinghosaurus from China. Acta Paleontologica Polonica, 50(4): 725~742. [PDF]

52.    *Wang X L, Zhou Z H, He H Y, Jin F, Wang Y Q, Zhang J Y, Wang Y, Xu X, Zhang F C, 2005. Stratigraphy and age of the Daohugou Bed in Ningcheng, Inner Mongolia. Chinese Science Bulletin, 50(20): 2369~2376.

 

2004

53.    *Wang Y, 2004a. A new Mesozoic caudate (Liaoxitriton daohugouensis sp. nov.) from Inner Mongolia, China. Chinese Science Bulletin (English edition), 49(8): 858~860. [PDF]

54.    *Wang Y, 2004b. Taxonomy and stratigraphy of late Mesozoic anurans and urodeles from China. Acta Geologica Sinica, 78(6): 1169~1178. [PDF]

55.    *Clark J M, Xu X, Forster C A, Wang Y, 2004. A Middle Jurassic ‘sphenosuchian’ from China and the origin of the crocodylian skull. Nature, 430: 1021~1024. [PDF]

 

2003

56.    Wang Y, Gao K Q, 2003. Amphibians. In: Chang M M, Chen P J, Wang Y Q, Wang Y (eds.), The Jehol Biota: The Emergence of Feathered Dinosaurs, Beaked Birds, and Flowering Plants. Shanghai: Shanghai Scientific & Techinial Publishers. 76~85. [pdf]

57.    Ting S Y, Bowen G J, Koch P L, Clyde W C, Wang Y Q, Wang Y, McKenna M C, 2003. Biostratigraphic, chemostratigraphic, and agnetostratigraphic study across the Paleocene-Eocene boundary in the Hengyang Basin Hunan, China. In: Wing S L, Gingerich P D, Schmitz B, Thomas E (eds), Causes and Consequences of Globally Warm Climates in the Early Paleogene: Boulder, Colorado: Geological Society of America Special, 369: 521~535.

58.    Chang M M, Chen P J, Wang Y Q, Wang Y (eds), 2003. The Jehol Biota: The Emergence of Feathered Dinosaurs, Beaked Birds and Flowering Plants. Shanghai: Shanghai Scientific & Technical Publishers. 208pp. // Chang M M, Chen P J, Wang Y Q, Wang Y, 2007. Atlas of the Jehol Biota Fossils: The Age of Feathered Dinosaurs. Asakura Publishing Co. Ltd. 202pp. (in Japanese) // Chang M M, Chen P J, Wang Y Q, Wang Y, 2008. The Jehol Fossils: The Emergence of Feathered Dinosaurs, Beaked Birds and Flowering Plants. Academic Press, Elsevier Ltd. 208pp.

2002

59.    Li C K, Wang Y (eds.), 2002. A Brief Introduction to Prehistoric Life. Beijing: Beijing Education Press. 180 pp. [in Chinese]

60.    *Swisher III C C, Wang X L, Zhou Z H, Wang Y Q, Jin F, Zhang J Y, Xu X, Zhang F C, Wang Y, 2002. Further Support for a Cretaceous age for the feathered-dinosaur beds of Liaoning, China: New 40Ar/39Ar dating of the Yixian and Tuchengzi Formations. Chinese Science Bulletin, 47(2): 135~138. [PDF]

61.    *Bowen G J, Clyde W C, Koch P L, Ting S Y, Alroy J, Tsubamoto T, Wang Y Q, Wang Y, 2002. Mammalian dispersal at the Paleocene/Eocene boundary. Science, 295: 2062~2065. [PDF]

62.    Zhang M M, Miao D S, Wang Y, 2002. Paleontological research in China: Retrospect and prospect. Bulletin of the Chinese Academy of Sciences, 16(2): 90~93. [PDF]

 

2001

63.    *Li J L, Wang Y, Wang Y Q, Li C K, 2001. A new family of primitive mammal from Liaoning, China. Chinese Science Bulletin (English Edition), 46(9): 782~786. [PDF]

64.    *Gao K Q, Wang Y, 2001. Mesozoic anurans from Liaoning Province, China, and phylogenetic relationships of archaeobatrachian anuran clades. Journal of Vertebrate Paleontology, 21(3): 460~476. [PDF]

65.    Wang Y, 2001. Advance in the study of Mesozoic lissamphibians from China. In: Deng T, Wang Y eds, Proceedings of the Eighth Annual Meeting of the Chinese Society of Vertebrate Paleontology. Beijing: China Ocean Press. 9~19. [in Chinese with English abstract] [PDF]

66.    Deng T, Wang Y (eds), 2001. Proceedings of the Eighth Annual Meeting of the Chinese Society of Vertebrate Paleontology. Beijing: China Ocean Press. 301 pp. [in Chinese with English abstract]

67.    Zhang M M, Chen P J, Wang Y Q, Wang Y (eds), 2001. The Jehol Biota. Shanghai: Shanghai Scientific & Technical Publishers. 150 pp. [in Chinese]

 

2000

68.    Wang Y, 2000. A new salamander (Amphibia: Caudata) from the Early Cretaceous Jehol Biota, Vertebrata PalAsiatica, 38(2): 100~103. [in Chinese with English abstract] [PDF]

69.    Wang X L, Wang Y Q, Zhang J Y, Jin F, Zhou Z H, Xu X, Zhang F C, Wang Y, Hu Y M, Lü J C, Gu G, 2000. New discovery of fossil vertebrates in the Jehol Group and discussion of the non-marine Jurassic-Cretaceous boundary in northern China. In: Editorial Committee of the Proceedings of the Third National Stratigraphical Conference of China (ed.), Proceedings of the Third National Stratigraphical Conference of China. Beijing: Geological Publishing House. 252~259. [in Chinese with English abstract] [pdf]

70.    Guo J W, Wang Y, Yang X A, 2000. A new early Eocene ctenodactyloid rodent (Rodentia, Mammalia) and the associated mammalian fossils from Danjiangkou, Hubei. Vertebrata PalAsiatica, 38(4): 303~313. [in Chinese with English abstract] [PDF]

71.    *Wang Y, Gao K Q, Xu X, 2000. Early evolution of discoglossid frogs: new evidence from the Mesozoic of China. Naturwissenschaften, 87(9): 417~420. [PDF]

72.    Wang Y, 2000. Archaic lissamphibians from China. In: Chinese Academy of Sciences (ed.), Innovators’ Report. Beijing: Science Press. 62~72. [in Chinese] [PDF]

 

1999

73.    *Swisher III C C, Wang Y Q, Wang X L, Xu X, Wang Y, 1999. Cretaceous age for the feathered dinosaurs of Liaoning, China. Nature. 400: 58~61. [PDF]

74.    *Wang Y, Gao K Q, 1999. Earliest Asian discoglossid frog from western Liaoning. Chinese Science Bulletin, 44(7): 636~642. [PDF] /

75.    Wang X L, Wang Y Q, Jin F, Xu X, Wang Y, 1999. Vertebrate assemblages of the Jehol biota in western Liaoning, China. In: Wang Y Q, Deng T (eds), Proceedings of the Seventh Annual Meeting of the Chinese Society of Vertebrate Paleontology. Beijing: China Ocean Press. 1~12. [pdf]

76.    Wang Y, Guo J W, Wang J W, 1999. A review of Chinese brontotheres. In: Wang Y Q, Deng T (eds), Proceedings of the Seventh Annual Meeting of the Chinese Society of Vertebrate Paleontology. Beijing: China Ocean Press. 139~147. [pdf]

77.    Wang X L, Wang Y Q, Xu X, Wang Y, Zhang J Y, Zhang F C, Jin F, Gu G, 1999. Record of the Sihetun vertebrate mass mortality events, western Liaoning, China: caused by volcanic eruptions. Geological Review, 45(supp): 458~467. [pdf]

78.    Wang X L, Wang Y Q, Jin F, Xu X, Wang Y, Zhang J Y, Zhang F C, Tang Z L, Li C, Gu G, 1999. The Sihetun fossil vertebrate assemblage and its geological setting of western Liaoning, China. PalaeoWorld. (11): 310~327. [pdf]

 

1998

79.    Wang X L, Wang Y Q, Wang Y, Xu X, Tang Z L, Zhang F C, Hu Y M, Gu G, Hao Z L, 1998. Stratigraphic sequence and vertebrate-bearing beds of the lower part of the Yixian Formation in Sihetun and neighboring area, western Liaoning, China. Vertebrata PalAsiatica, 36(2): 81~101. [PDF]

80.    Dong Z M, Wang Y, 1998. A new urodele (Liaoxitriton zhongjiani gen. et sp. nov.) from the Early Cretaceous of western Liaoning Province, China. Vertebrata PalAsiatica, 36(2): 159~172. [PDF]

 

1997

81.    Wang Y, Wang J W, 1997. A new brontothere from late middle Eocene of Qufu, Shandong. Vertebrata PalAsiatica, 35(1): 68~77. [PDF]

 

1995

82.    Wang Y, 1995. A new primitive chalicothere (Perissodactyla, Mammalia) from the early Eocene of Hubei, China. Vertebrata PalAsiatica, 33(2): 138~159. [PDF]

 

 

 

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